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- First evidence of a non-thalattosuchian mesoeucrocodylian from the Upper Jurassic of Spain (Villar del Arzobispo Formation, Alpuente)Publication . Sánchez Fenollosa, Sergio; Suñer, Maite; Gamonal, ArturoABSTRACT: Non-thalattosuchian mesoeucrocodylian fossil remains from the Upper Jurassic of Europe are mainly comprised by isolated teeth, and come from Portugal, France and Germany. In the present study, we report the occurrence of a small crocodylomorph tooth from a new fossil site named “Puente de La Hortichuela” located in the Villar del Arzobispo Formation on the municipality of Alpuente (Valencia, Spain). The tooth has been recovered from the lower third of this lithostratigraphic unit with an Upper Jurassic age. The tooth has molariform or tribodont morphology, kidney-shape cross section, and enamel ornamentation formed by parallel basiapical ridges. These characteristics lead us to tentatively identify the tooth as belonging to the middle or posterior dentition of a bernissartiid. This finding is the oldest record of a non-thalattosuchian mesoeucrocodylian from Spain and provides new information about the fauna that inhabited this region.